export do SBML throws error 500m; cannot correct aliases in CellD file

Attached file throws below error while export as SBML:
{"reason":"[Complex csa24]\tProblem with exporting bioEntity","error":"Internal server error."} I tried to re-draw the map, to change/update aliases but the error is reoccurring. It is the same issue like #1756 (closed), but in #1756 (closed) I was able to fix incorrect alias, while in this example - I cannot. Axonal-remodeling-and-CDK5-signaling.xml